The Silent Shrimp Slower

How a Tiny Parasite is Disrupting India's Seafood Treasure

Aquaculture EHP Shrimp Farming West Bengal

Introduction

Imagine a shrimp farmer in West Bengal, patiently tending to his ponds for months, anticipating a healthy harvest of Pacific White shrimp. As the harvest day approaches, he notices something puzzling—his shrimp show extreme size variations, with many remaining disappointingly small despite months of feeding.

The mystery culprit behind this economic tragedy isn't a virus or bacterium, but a microscopic parasite called Enterocytozoon hepatopenaei (EHP), silently wreaking havoc in aquaculture ponds across Eastern India 1 2 .

This article explores the fascinating detective story of how scientists identified and characterized this emerging threat to India's vital shrimp industry, focusing on groundbreaking research from West Bengal that revealed one of the highest infection rates ever documented in the country 1 .

Size Variation

Extreme differences in shrimp size within the same pond

Growth Retardation

Shrimp remain small despite adequate feeding

Economic Loss

Undersized shrimp command lower market prices

Meet the Culprit: Ecytonucleospora hepatopenaei

EHP (Ecytonucleospora hepatopenaei, previously known as Enterocytozoon hepatopenaei) is a microsporidian parasite—a type of fungal-related, spore-forming organism that can only survive and reproduce inside the cells of its host 5 8 .

The parasite primarily infects the hepatopancreas of shrimp, a crucial digestive organ that functions similarly to the human liver and pancreas combined 6 . This organ is responsible for producing digestive enzymes, absorbing nutrients, and storing energy reserves—all essential processes for growth and development 6 .

EHP was first reported in black tiger shrimp in Thailand in 2004 6 8 but has since spread to numerous shrimp-farming countries across Asia. The parasite made its way to India with the introduction of the non-native Pacific White shrimp (Litopenaeus vannamei) in 2009, which revolutionized Indian aquaculture but inadvertently introduced new disease challenges 4 .

Microsporidian Parasite

Fungal-related, spore-forming intracellular parasite

Hepatopancreas Target

Primary infection site - the shrimp's digestive organ

The West Bengal Outbreak: Ground Zero for EHP

In 2019, scientists conducted a comprehensive investigation into the growing reports of stunted shrimp across West Bengal's coastal districts 1 . What they discovered was alarming—EHP had firmly established itself throughout the region with devastating prevalence.

EHP Prevalence in West Bengal's Shrimp Farming Districts

District Samples Collected Infection Rate
East Midnapur 119 samples 84.9% overall prevalence
North 24 Parganas 50 samples Part of overall 84.9% prevalence
South 24 Parganas 50 samples Part of overall 84.9% prevalence

84.9%

Overall EHP Prevalence Rate

Highest reported in India at the time 1

The Detective Work: How Scientists Uncover EHP

Step 1: Recognizing the Signs

Shrimp farmers first noticed concerning signs in their ponds:

  • Extreme size variation—a mix of normal and stunted shrimp in the same pond
  • Floating white fecal strings on the pond surface
  • Reduced feeding activity
  • Soft shells and lethargic behavior
  • Empty midguts in affected shrimp 2 6
Field Observations

Farmers report unusual patterns in shrimp growth and behavior that signal potential EHP infection.

Step 2: Laboratory Confirmation

When these symptomatic shrimp arrived at laboratories, scientists employed multiple techniques to confirm EHP infection:

Histopathological Analysis

Researchers examined thin sections of hepatopancreas tissue under the microscope, revealing telltale signs of EHP infection—eosinophilic to basophilic inclusions within the epithelial cells of the hepatopancreatic tubules, along with moderate necrotic tubular detachment from the basal membrane 1 7 .

Molecular Detection with PCR

The most definitive diagnosis came from polymerase chain reaction (PCR) testing. Scientists extracted DNA from hepatopancreatic tissues and used specialized primers targeting two different EHP genes:

  • ssu rRNA gene: Produced expected PCR products of 951 and 510 bp
  • Spore wall protein (SWP) gene: A second-generation primer providing even more specific detection 1
Molecular Detection Methods for EHP
Detection Method Target Gene Amplicon Size
Conventional PCR ssu rRNA 951 bp and 510 bp
Second-generation PCR Spore wall protein (SWP) Varies
Nested PCR ssu rRNA and SWP 514 bp and 148 bp

Scientific Toolkit: Essential Research Tools

Modern EHP research relies on sophisticated laboratory tools and reagents:

Research Tool Specific Example Function in EHP Research
DNA Extraction Kits DNeasy Blood & Tissue kits Isolate high-quality DNA from shrimp tissues
PCR Primers SWP_1F/SWP_1R and SWP_2F/SWP_2R Amplify specific EHP gene sequences for detection
Histology Reagents Davidson's AFA solution Fix and preserve tissue for microscopic examination
Staining Agents Hematoxylin and Eosin (H&E) Visualize tissue structure and pathological changes
Molecular Biology Reagents Taq DNA polymerase, dNTPs, MgCl2 Enable DNA amplification through PCR
Genetic Analysis

Genetic sequencing of PCR products revealed 100% homology with EHP sequences from other affected countries including Vietnam, Thailand, China, and other parts of India 1 .

Tissue Examination

Histopathological analysis provides visual confirmation of EHP infection through characteristic cellular changes in hepatopancreatic tissues.

The Domino Effect: How a Tiny Parasite Causes Widespread Damage

Broken Digestion

The hepatopancreas, as the primary digestive organ, becomes increasingly compromised as EHP multiplies within its cells. The parasite causes:

Reduced Enzyme Production

Impaired digestive enzyme synthesis

Nutrient Malabsorption

Impaired nutrient uptake

Tubular Atrophy

Wasting away of hepatopancreatic tubules

Cellular Necrosis

Death of tubular epithelial cells 6 7

With their digestive system compromised, infected shrimp cannot effectively convert feed into growth, leading to the characteristic stunting despite adequate feeding.

Beyond Growth: The White Feces Connection

Researchers also noted an association between EHP infection and White Feces Syndrome (WFS), a condition where shrimp produce white, stringy fecal matter that floats on the pond surface 1 7 . Studies confirmed that these floating white feces contained EHP spores, creating an efficient transmission route within ponds 7 .

White Feces Syndrome

Floating white fecal strings containing EHP spores serve as transmission vectors.

Beyond Growth: Secondary Infections and Microbiome Disruptions

Microbiome Mayhem

A 2022 study revealed that EHP infection follows a degenerative cyclic pattern in the hepatopancreas microbiome 9 . In early stages, high diversity of beneficial microorganisms related to nutrient assimilation persists. As the infection progresses, these beneficial organisms decrease while opportunistic pathogenic fungi increase. In late stages, opportunistic bacteria and fungi associated with disease processes displace the beneficial microorganisms 9 .

Immune System Dysregulation

Transcription analysis revealed significant changes in immune gene expression:

  • Upregulated genes: ProPO, Hsp70, and α2-macroglobulin
  • Downregulated genes: LGBP, PXN, and Integrin ß 6
Secondary Infections

Gut samples from symptomatic shrimp contained pathogenic V. parahaemolyticus, V. harveyi, and V. campbellii—all known to cause serious diseases in shrimp 6 . This helps explain why EHP-infected shrimp often succumb to secondary bacterial infections.

Fighting Back: Prevention and Control

Currently, no approved therapeutics exist to treat EHP infections in shrimp 5 . Control strategies therefore focus on prevention and management.

Biosecurity

Preventing introduction of EHP through PCR-screened, EHP-free postlarvae 5

Disinfection

Proper pond preparation between cycles to eliminate spores from previous crops 5

Management

Avoiding practices that promote EHP transmission, such as collecting floating white feces 7

Probiotics

Exploring microbiome manipulation to enhance shrimp health and resistance to EHP 9

Research Directions

Ongoing research continues to unravel the complex interactions between EHP and its shrimp host, particularly the immune response triggered by infection 8 and the disruption of the gut microbiome 9 . Each new discovery provides potential avenues for future management strategies.

Conclusion: A Continuing Battle

The story of EHP in West Bengal's shrimp farms represents a classic case of an emerging pathogen finding opportunity in a rapidly expanding industry. The 84.9% prevalence rate discovered in 2019 served as a wake-up call to the region's aquaculture sector 1 .

While EHP doesn't typically cause mass mortality like some viral diseases, its impact on growth and size uniformity creates substantial economic losses for farmers who must harvest undersized shrimp that command lower market prices 4 5 .

As the global demand for seafood continues to grow and aquaculture expands, understanding and managing microscopic threats like EHP will become increasingly important—not just for West Bengal, but for seafood production worldwide.

Global Aquaculture Challenge

References

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References